Lawrence W. Sheppard

Postdoctoral Researcher
Primary office:


Faculty Sponsor:  Daniel Reuman

Selected Publications




Area of specialization
Wavelet analysis (data analysis of time-varying signals, with biological, physical or ecological applications)

Research interests:

  • Statistical analysis of wavelet coherence
  • Intracranial pressure analysis (biomedical physics application)
  • Skin temperature, anesthesia and ensembles of neurons (biomedical physics application)
  • Aphid monitoring and phenology
  • Plankton monitoring and climate

Selected Publications:

Anderson, T.L., Sheppard, L.W., Walter, J.A., Hendricks, S.P., Levine, T.L., White, D.S., Reuman, D.C.. 2019. The dependence of synchrony on timescale and geography in freshwater plankton. Limnology and Oceanography. 64: 483-502. doi: 10.1002/lno.11054.

Zhao., L., Sheppard, L.W., Reid, P.C., Walter, J.A., and Reuman, D.C.. 2019. Proximate determinants of Taylor’s law slopes. Journal of Animal Ecology 88, 484-494. doi: 10.1111/1365-2656.12931.

Sheppard, L.W., Defriez, E.J., Reid, P.C., Reuman, D.C. 2019. Synchrony is more than its top-down and climatic parts: interacting Moran effects on phytoplankton in British seas. PloS Computational Biology 15: e1006744. doi: 10.1371/journal.pcbi.1006744.

Walter, J.A., Sheppard, L.W., Venugopal, P.D., Reuman, D.C., Dively, G., Tooker, J.F., Johnson, D.M. 2019. Weather and regional crop composition variation drive spatial synchrony of lepidopteran agricultural pests. Ecological Entomology. doi: 10.1111/een.12830.

Sheppard, L.W., Zhao, L., Anderson, T.L., Walter, J.A., and Reuman, D.C. 2018.  wsyn: Wavelet approaches to synchrony.  A software package for the R programming language, published after review on the Comprehensive R Archive Network (CRAN).  Developmnt version at

Reuman, D., Zhao, L., Sheppard, L.W., Reid, P.C. Cohen, J.E. 2018.  Synchrony affects Taylor's law in theory and data.  Proceedings of the National Academy of Science 114: 6788-6793.

Walter, J.A., Sheppard, L.W., Anderson, T.L., Kastens, J.H., Bejornstad, O.N., Liebhold, A.M., and Reuman, D.C. 2017.  The geography of synchrony.  Ecology Letters 20: 801-814.

Sheppard, L.W., Reid, P.C., and Reuman, D.C. 2017.  Rapid surrogate testing of wavelet coherences.  European Physical Journal, Nonlinear and Biomedical Physics 5:1.

Reuman, D.C., Zhao, L., Sheppard, L.W., Reid, P.C., and Cohen, J.E. 2017.  Synchrony affects Taylor's law in theory and data.  Proceedings of the National Academy of Sciences 114: 6788-6793.

defriez, E., Sheppard, L., Reid, P.C., and Reuman, D.C.  2016.  Climate-change-related regime shifts have altered spatial synchrony of plankton dynmaics in the North Sea.  Global Change Biology 10: 1111/gcb.13229.

Sheppard, L., Bell, J., Harrington, R., and Reuman, D.  2015.  Changes in large-scale climate alter spatial synchrony of aphid pests.  Nature Climate Change 6: 610-613. 

Recent Presentations:

  • Ghosh, S., Hallett, L.M., Sheppard, L.W., and Reuman, D. 2019. Estimating synchronous and compensatory dynamics in community ecology using a novel “copula” approach.  Ecological Society of America, Louisville, KY., August



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