Rafe Marion Brown

College of Liberal Arts and Sciences - Ecology & Evolutionary Biology, KU Biodiversity Institute
Associate Professor
Curator-in-Charge, Herpetology Division, Natural History Museum and Biodiversity Institute
Primary office:
785-864-3403
Dyche Hall
Room 417
Dyche Hall, 1345 Jayhawk Blvd.
University of Kansas
Lawrence, KS 66045
Second office:



Dr. Rafe Brown is a herpetologist interested in understanding evolutionary processes of diversification in island archipelagos.

Teaching Interests

  • Herpetology
  • Biogeography
  • NSF DDIG Grant Writing

Research

Using molecular markers in conjunction with statistical approaches to phylogeny estimation, I am interested in testing hypotheses regarding species boundaries, distributions, dispersal, isolation, and diversity. Questions of lineage diversification and species-specific responses to the geographic template are particularly attractive research topics for an empirical systematist working in island archipelagos. My fieldwork is based primarily in the archipelagos of the Philippines, Indonesia, and the Solomon Islands where I take a comparative, multi-taxon approach to biogeographical studies of land vertebrates. As a systematic herpetologist, I maintain a commitment to biodiversity studies, taxonomy, and conservation of amphibians and reptiles of Southeast Asia.

Research Interests

  • Biogeography and processes of diversification in island archipelagos
  • Phylogenetic systematics
  • Character evolution
  • Speciation genetics
  • Biodiversity, conservation and taxonomy of amphibians and reptiles of Southeast Asia

Selected Publications

Brown, R. M. (in press). A new species of stream frog (genus Hylarana) from the mountains of southern Mindanao Island, Philippines. Herpetologica

Brown, R. M., Richards, S. J., & Broadhead, T. (in press). The first record of the genus Batrachylodes (Anura: Ceratobatrachidae) outside of the Solomon Islands archipelago, and a description of a new species from New Britain. Zootaxa

Wynn, A., Diesmos, A. C., & Brown, R. M. (in press). Two new species of Malayotyphlops from the northern Philippines, with redescriptions of M. luzonensis (Taylor) and M. ruber (Boettger). Journal of Herpetology

Barley, A. J., Monnahan, P. J., Thomson, R., Grismer, L. L., & Brown, R. M. (2015). Sun skink landscape genomics across Southeast Asia: the evolution of genetic and morphological diversity across a heterogeneous and fragmented landscape. Molecular Ecology, 1696–1712.

Brown, R. M., Siler, C. D., Richards, S., Diesmos, A. C., & Cannatella, D. C. (2015). Multilocus phylogeny and a new classification for Southeast Asian and Melanesian forest frogs (family Ceratobatrachidae). Zoological Journal of the Linnaean Society.

Diesmos, A. C., Alcala, A. C., Siler, C. D., & Brown, R. M. (2014). Status and Conservation of Philippine Amphibians. In H. Heatwole & I. Das (Eds.), Conservation Biology of Amphibians of Asia. Status of Conservation and Decline of Amphibians: Eastern Hemisphere (pp. 310-336). Kota Kinabalu, Malaysia: Natural History Publications (Borneo).

Leviton, A. E., Brown, R. M., & Siler, C. D. (2014). The dangerously venomous snakes of the Philippine archipelago, with identification keys and species accounts. In G. C. Williams & G. C. Gosliner (Eds.), The Coral Triangle: the 2011 Hearts Philippine Biodiversity Expedition (pp. 473-530). San Francisco, CA: California Academy of Sciences.

Chan, K. O., Brown, R. M., Ahmad, N., & Grismer, L. Lee (2014). A new species of frog (Amphibia: Anura: Ranidae) of the Hylarana signat Complex from Peninsular Malaysia. Herpetologica, 70, 228–240.

Gonzalez, P., Yu, Y., Siler, C. D., Barley, A., Sanguila, M. B., Diesmos, A. C., & Brown, R. M. (2014). Archipelago colonization by ecologically dissimilar amphibians: evaluating the expectation of common evolutionary history of geographical diffusion in co-distributed rainforest tree frogs in islands of Southeast Asia. Molecular Phylogenetics and Evolution, 72, 35-41.

Siler, C. D., Lira-Noriega, A., & Brown, R. M. (2014). Conservation genetics of Australasian Sailfin Lizards: flagship species threatened by coastal development and insufficient protected area coverage. Biological Conservation, 169, 100-108.

Brown, R. M., Weghorst, J. A., Olson, K. V., Duya, M. R. M., Barley, A. J., Duya, L. V., Shekelle, M., Neri-Arboleda, I., Esselstyn, J. A., Dominy, N. J., Ong, P. S., Moritz, G. L., Luczon, A., Diesmos, M. L. D., Diesmos, A. C., & Siler, C. D. (2014). Conservation genetics of the Philippine tarsier: cryptic genetic variation restructures conservation priorities for an island archipelago primate. PLoS ONE, 9, e104340. DOI:10.1371/journal.pone.0104340

Siler, C. D., Oaks, J. R., Cobb, K., Ota, H., & Brown, R. M. (2014). Critically endangered island endemic or peripheral population of a widespread species? Conservation genetics of Kikuchi’s gecko and the global challenge of protecting peripheral oceanic island endemic vertebrates. Diversity and Distributions, 20, 756–772.

Siler, C. D., Dececchi, A. A., Merkord, C. L., Davis, D. R., Christiani, T. J., & Brown, R. M. (2014). Cryptic diversity and population genetic structure in the rare, endemic, forest-obligate, forest geckos of the Philippines. Molecular Phylogenetics and Evolution, 70, 204–209.

Davis, D., Feller, K. D., Brown, R. M., & Siler, C. D. (2014). Evaluating the diversity of Philippine Slender Skinks of the Brachymeles bonitae Complex (Reptilia: Squamata: Scincidae): Redescription of B. tridactylus and descriptions of two new species. Journal of Herpetology, 480–494.

Welton, L. J., Wood, P. L., Oaks, J. R., Siler, C. D., & Brown, R. M. (2014). Fossil-calibrated phylogeny and historical biogeography of Southeast Asian water monitors (Varanus salvator Complex). Molecular Phylogenetics and Evolution, 74, 29–37.

Welton, L. J., Siler, C. D., Travers, S. N., & Brown, R. M. (2014). Integrative taxonomy and phylogeny-based species delimitation of Philippine water monitor lizards (Varanus salvator Complex) with descriptions of two new cryptic species. Zootaxa, 3881, 201–227.

Chan, K. O., Grismer, L. L., & Brown, R. M. (2014). Reappraisal of the Javanese bullfrog complex, Kaloula baleata (Müller, 1836) (Amphibia: Anura: Microhylidae), reveals a new species from northwestern Peninsular Malaysia. Zootaxa, 3900, 569–580.

Rocha, L. A., Aleixo, A., Allen, G., Almeda, F., Baldwin, C., Barclay, M., Bates, J., Bauer, A., Benzoni, F., Berns, C., Berumen, M., Blackburn, D. C., Blum, S., Bolaños, F., Bowie, R., Brit, R., & Brown, R. M. (2014). Specimen collection: an essential tool. Science, 344, 814–815.

Barley, A. J., Datta-Roy, A., Karanth, K. P., Thompson, R., & Brown, R. M. (2014). Sun skink diversification across the Indian-Southeast Asian biogeographical interface. Journal of Biogeography. DOI:10.1111/jbi.12397

Siler, C. D., Welton, L. J., Davis, D. R., Watters, J. L., Davey, C. S., Diesmos, A. C., Diesmos, M. L., & Brown, R. M. (2014). Taxonomic revision of the Pseudogekko compresicorpus Complex (Reptilia: Squamata: Gekkonidae), with descriptions of three new species. Herpetological Monographs, 28, 110–139.

Siler, C. D., Linkem, C. W., Cobb, K., Watters, J. L., Cumings, S., Diesmos, A. C., & Brown, R. M. (2014). Taxonomic revision of the semi-aquatic skink Parvoscincus leucospilos (Reptilia: Squamata: Scincidae), with description of three new species. Zootaxa, 3847, 388–412.

Richards, S. J., Oliver, P., & Brown, R. M. (2014). Monograph Series Vol. 2. A new scansorial species of Platymantis Günther 1858 (Anura: Ceratobatrachidae) from Manus Island, Admiralty Archipelago, Papua New Guinea. Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, 123–134. Lettland, Latvia: The Entomological Society of Latvia.

Brown, R. M., Richards, S. J., & Broadhead, T. S. (2013). A new shrubfrog in the genus Platymantis (Ceratobatrachidae) from the Nakanai Mountains of eastern New Britain Island, Bismarck Archipelago. Zootaxa, 3710, 31-45.

Blackburn, D. C., Siler, C. D., Diesmos, A. C., McGuire, J. A., Cannatella, D. C., & Brown, R. M. (2013). An adaptive radiation of frogs in a Southeast Asian island archipelago. Evolution.

Welton, L. J., Siler, C. D., Linkem, C. W., Diesmos, A. C., Diesmos, M. L., Sy, E., & Brown, R. M. (2013). Dragons in our midst: phyloforensics of illegally traded Southeast Asian water monitor lizards. Biological Conservation, 159, 7-15.

Oaks, J. R., Sukumaran, J., Esselstyn, J. A., Linkem, C. W., Siler, C. D., Holder, M. T., & Brown, R. M. (2013). Evidence for Pleistocene-driven diversification? A caution for interpreting ABC inferences of simultaneous historical events. Evolution, 67, 991–1010.

Brown, R. M., Siler, C. D., Oliveros, C. H., Esselstyn, J. A., Diesmos, A. C., Hosner, P. A., Linkem, C. W., Barley, A. J., Oaks, J. R., Sanguila, M. B., Welton, L. J., Moyle, R. G., Peterson, A. T., & Alcala, A. C. (2013). Evolutionary processes of diversification in a model island archipelago. Annual Review of Ecology, Evolution, and Systematics, 44, 411–435.

Grismer, L. L., Wood, P. L., Anuar, S., Muin, M. A., Quah, E. S. H., McGuire, J. A., Brown, R. M., & Tri, N. V. (2013). Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia. Zoological Journal of the Linnean Society, 169, 849–880.

Welton, L. J., Siler, C. D., Oaks, J. R., Diesmos, A. C., & Brown, R. M. (2013). Multilocus phylogeny and Bayesian estimates of species boundaries reveal hidden evolutionary relationships and cryptic diversity in Southeast Asian monitor lizards. Molecular Ecology, 22, 3495–3510.

Siler, C. S., Oliveros, C. H., Santannen, A., & Brown, R. M. (2013). Multilocus phylogeny reveals unexpected patterns of regional and morphological diversification in Asian Wolf Snakes (genus Lycodon). Zoologica Scripta.

Chan, K. O., Blackburn, D. C., Murphy, R. W., Stuart, B. L., Emmett, D. A., Thu Ho, C., & Brown, R. M. (2013). New species of narrow-mouthed frog of the genus Kaloula from eastern Indochina. Herpetologica, 69, 329–341.

Siler, C. D., Welton, L. J., Rock, A. A., & Brown, R. M. (2013). Population genetic structure and revised geographic range for the tridactyl skink (Brachymeles muntingkamay) from Luzon Island, Philippines. Herpetological Review, 44, 30-33.

Brown, R. M., & Siler, C. D. (2013). Spotted stream frog diversification at the Australasian faunal zone interface, mainland versus island comparisons, and a test of the Philippine ‘dual-umbilicus’ hypothesis. Journal of Biogeography, 41, 182–195.

Linkem, C. W., Brown, R. M., Siler, C. D., Evans, B. J., Austin, C. C., Iskandar, D. T., Diesmos, A. C., Supriatna, J., Andayani, N., & McGuire, J. A. (2013). Stochastic faunal exchanges drive diversification in widespread Wallacean and Pacific Island lizards (Squamata: Scincidae: Lamprolepis smaragdina). Journal of Biogeography, 40, 507–520.

Linkem, C. W., & Brown, R. M. (2013). Systematic revision of the Parvoscincus decipiens (Boulenger 1894) complex of Philippine forest skinks (Squamata: Scincidae: Lygosominae) with descriptions of seven new species. Zootaxa, 3700, 501–533.

Brown, R. M., Siler, C. D., Oliveros, C. H., Welton, L. J., Rock, A., Swab, J., Van Weerd, M., van Beijnen, J., Jose, E., Rodriguez, D., Jose, E., & Diesmos, A. C. (2013). The amphibians and reptiles of Luzon Island, Philippines, VIII: the herpetofauna of Cagayan and Isabela Provinces, northern Sierra Madre Mountain Range. Zookeys, 266, 1-120.

Barley, A., White, J., Diesmos, A. C., & Brown, R. M. (2013). The challenge of species delimitation at the extremes: diversification without morphological change in Philippine sun skinks. Evolution, 67, 3556–3572.

Böhm, M., Collen, B., Baillie, J. E. M., Chanson, J., Cox, N., Hammerson, G., Hoffmann, M., Livingstone, S. R., Rhodin, A. G. J., Stuart, S. N., van Dijk, P. Pl., Young, B. E., Afuang, L. E., Aghasyan, A., Aguayo, A. G., Aguilar, C., Akarsu, F., Alencar, L. R. V., Allison, A., Ananjeva, N., Anderson, S., Andrén, C., Ariano-Sánchez, D., Arrendondo, J. C., Auliya, M., Austin, C. C., Avci, A., Baker, P. J., Barreto-Lima, A. F., Barrio-Amorós, C. L., Basu, D., Bates, M. F., Batistella, A., Bauer, A., Bennett, D., Böhme, W., Broadley, D., Brown, R., Burgess, J., Captain, A., Carreira, S., Castaneda, M., Castro, F., Catenazzi, A., Cedeño-Vázquez, J. R., Chapple, D., Cheylan, M., Cisneros-Heredia, D. F., Cogalniceanu, D., Cogger, H., Corti, C., Costa, G. C., Couper, P. J., Courtney, T., Crnobrnja-Isailovic, J., Crochet, P., Crother, B., Cruz, F., Daltry, J., Daniels, R. J. R., Das, I., de Silva, A., Dirksen, L., Doan, T., Dodd, K., Doody, J. S., Dorcas, M. E., Duarte de Barros Filho, J., Egan, V. T., El Mouden, E. H., Embert, D., Espinoza, R. E., Fallabrino, A., Feng, X., Feng, Z., Fitzgerald, L., Flores-Villela, O., França, F. G. R., Frost, D., Gadsden, H., Gamble, T., Ganesh, S. R., Garcia, M. A., García-Pérez, J. E., Gatus, J., Gaulke, M., Geniez, P., Georges, A., Gerlach, J., Goldberg, S., Gonzalez, J. C. T., Gower, D. J., Grant, T., Greenbaum, E., Guo, P., Haitao, S., Hamilton, A. M., Hare, K., Hedges, B., Heideman, N., Hilton-Taylor, C., Hitchmough, R., Hollingsworth, B., Hutchinson, M., Ineich, I., Iverson, J., Jaksic, F. M., Jenkins, R., Joger, U., Jose, R., Kaska, Y., Keogh, J. S., Köhler, G., Kuchling, G., Kumlutas, Y., Kwet, A., La Marca, E., Lamar, W., Lane, A., Lardner, B., Latta, C., Latta, G., Lau, M., Lavin, P., Lawson, D., LeBreton, M., Lehr, E., Limpus, D., Lipczynski, N., Lobo, A. S., López-Luna, M. A., Luiselli, L., Lukoschek, V., Lundberg, M., Lymberakis, P., Macey, R., Magnusson, W. E., Mahler, L., Malhotra, A., Mariaux, J., Maritz, B., Marques, O. A. V., Márquez, R., Martins, M., Masterson, G., Mateo, J. A., Mathew, R., Mathews, N., Mayer, G., McCranie, J. R., Measey, J., Mendoza-Quijano, F., Menegon, M., Métrailler, S., Milton, D. A., Montgomery, C., Morato, S. A. A., Mott, T., Muñoz-Alonso, A., Murphy, J., Nguyen, T. Q., Nilson, G., Nogueira, C., Núñez, H., Ota, H., Ottenwalder, J., Papenfuss, T., Pasachnik, S., Passos, P., Pauwels, O. S. G., Pérez Mellado, V., Pérez-Buitrago, N., Pianka, E. R., Pleguezuelos, J., Pollock, C., Ponce-Campos, P., Powell, R., Pupin, F., Quintero Díaz, G. E., Radder, R., Ramer, J., Rasmussen, A. R., Raxworthy, C., Reynolds, R., Richman, N., Rico, E. L., Riservato, E., Rivas, G., Rocha, P. L. B., Rödel, M., Rodríguez Schettino, L., Roosenburg, W. M., Ross, J. P., Sadek, R., Sanders, K., Santos-Barrera, G., Schleich, H. H., Schmidt, B., Schmitz, A., Sharifi, M., Shea, G., Shine, R., Slimani, T., Somaweera, R., Spawls, S., Stafford, P., Stuebing, R., Sweet, S., Sy, E., Temple, H., Tognielli, M., Tolley, K., Tolson, P. J., Tuniyev, B., Tuniyev, S., Üzüm, N., van Buurt, G., Van Sluys, M., Velasco, A., Vences, M., Veselý, M., Vinke, S., Vinke, T., Vogel, G., Vogrin, M., Vogt, R. C., Wearn, O. R., Werner, Y. L., Whiting, M. J., Wiewandt, T., Wilkinson, J., Wilson, B., Wren, S., Zamin, T., Zhou, K., & Zug, G. (2013). The conservation status of the world’s reptiles. Biological Conservation, 157, 372–385.

Brown, R. M., & Stuart, B. L. (2012). Patterns of biodiversity discovery through time: an historical analysis of amphibian species discoveries in the Southeast Asian mainland and island archipelagos. In D. J. Gower, K. G. Johnson, J. E. Richardson, B. R. Rosen, L. Rüber, & S. T. Williams (Eds.), Biotic Evolution and Environmental Change in Southeast Asia (pp. 348-389). Cambridge University Press.

Brown, R. M. (2012). [Review of the book Biogeography 4th Edition, Lomolino, Riddle & Brown (Sinauer)]. Quarterly Review of Biology, 87, 153-154.

Brown, R. M., Diesmos, A. C., Sanguila, M. B., Siler, C. D., Diesmos, M. L. D., & Alcala, A. C. (2012). Amphibian conservation in the Philippines. FrogLog, 104, 40-43.

Brown, R. M., Oliveros, C. H., Siler, C. D., Fernandez, J. B., Welton, L. J., Buenavente, P. A. C., Diesmos, M. L. D., & Diesmos, A. C. (2012). Amphibians and Reptiles of Luzon Island (Philippines), VII: Herpetofauna of Ilocos Norte Province, Northern Cordillera Mountain Range. Check List, 8, 469-490.

Devan-Song, A., & Brown, R. M. (2012). Amphibians and Reptiles of Luzon Island, Philippines, VI: the Herpetofauna of the Subic Bay Area. Asian Herpetological Research, 2012, 1-20.

Siler, C. D., Swab, J. C., Oliveros, C. H., Diesmos, A. C., Averia, L., Alcala, A. C., & Brown, R. M. (2012). Amphibians and reptiles, Romblon Island Group, central Philippines: comprehensive herpetofaunal inventory. Check List, 8, 443-462.

Spinks, P. Q., Thomson, R. C., Hughes, B., Moxley, B., Brown, R. M., Diesmos, A. C., & Shaffer, H. B. (2012). Cryptic variation and the tragedy of unrecognized taxa: the case of international trade in the spiny turtle Heosemys spinosa (Testudines: Geoemydidae). Zoological Journal of the Linnean Society, 164, 811-824.

Alcala, A. C., Bucol, A. A., Alcala, E. L., & Brown, R. M. (2012). Decreasing population size of the Philippine Limestone Frog, Platymantis insulatus. FrogLog, 104, 44-45.

Diesmos, M. L. D., Diesmos, A. C., Siler, C. D., Vredenburg, V. T., & Brown, R. M. (2012). Detecting the Distribution of the Chytrid Fungus in the Philippines. FrogLog, 104, 48-49.

Siler, C. D., Oaks, J. R., Welton, L. J., Linkem, C. W., Swab, J., Diesmos, A. C., & Brown, R. M. (2012). Did geckos ride the Palawan raft to the Philippines? Journal of Biogeography, 39, 1217-1234.

Welton, L. J., Siler, C. D., Diesmos, A. C., Diesmos, M. L. D., Lagat, R., Causaren, R., & Brown, R. M. (2012). Genetic identity, geographic range, and major distribution records for frugivorous monitor lizards of Luzon Island, Philippines. Herpetological Review, 43, 226-230.

Siler, C. D., Oliveros, C. H., & Brown, R. M. (2012). Geographic range notes on Brachymeles cebuensis and re-evaluation of its conservation status. Herpetological Review, 43, 305.

Brown, R. M., Siler, C. D., Grismer, L. L., Das, I., & McGuire, J. A. (2012). Phylogeny and cryptic diversification in Southeast Asian flying geckos. Molecular Phylogenetics and Evolution, 65, 351-361.

Siler, C. D., Jones, R. M., Diesmos, A. C., Diesmos, M. L., & Brown, R. M. (2012). Phylogeny-based species delimitation in Philippine slender skinks (Reptilia: Squamata: Scincidae) III: taxonomic revision of the Brachymeles gracilis complex, with description of three new species. Herpetological Monographs, 26, 135-172.

Brown, R. M., Siler, C. D., Das, I., & Min, P. Y. (2012). Testing the phylogenetic affinities of Southeast Asia’s rarest geckos: flap-legged geckos (Luperosaurus), flying geckos (Ptychozoon) and their relationship to the pan-Asian genus Gekko. Molecular Phylogenetics and Evolution, 63, 915-921.

Alcala, A. C., Bucol, A., Diesmos, A. C., & Brown, R. M. (2012). Vulnerability of Philippine amphibians to climate change. Philippine Journal of Science, 141, 77-87.

Diesmos, A. C., Buskirk, J. R., Schoppe, S., Diesmos, M. L. L., Sy, E. Y., & Brown, R. M. (2012). Che- lonian Research Monographs No. 5. Siebenrockiella leytensis (Taylor 1920) – Palawan Forest Turtle, Philippine Forest Turtle (A. G. J. Rhodin, P. C. H. Pritchard, P. P. van Dijk, R. A. Saumure, K. A. Buhlmann, J. B. Iverson, & R. A. Mittermeier). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group, 066.1–066.9. DOI:10.3854/ crm.5.066.leytensis.v1.2012 http://www.iucn-tftsg.org/cbftt/

Brown, R. M. (2012). Biodiversity information products for Philippine conservation. http://www.nhm.ku.edu/rbrown/PublicationsMain.htm

Siler, C. D., & Brown, R. M. (2012). Philippines Biodiversity Research Education and Outreach (PhilBREO) lifedesks. Encyclopedia of Life. http://philbreo.lifedesks.org/

Diesmos, A. C., & Brown, R. M. (2011). Diversity, Biogeography, and Conservation of Philippine Amphibians. In Biology and Conservation of Tropical Asian Amphibians. Proceedings of the Conference “Biology of the Amphibians in the Sunda Region, South-east Asia" (pp. 26-49). Kota Samarahan, Sarawak, Malaysia: Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak.

Phenix, E., Phenix, J. W., Siler, C., Brown, R. M., & Diesmos, A. C. (2011). Hologerrhum philipinum—reproductive mode. Herpetological Review, 43, 614.

Siler, C. D., Welton, L. J., Brown, R. M., Infante, C., & Diesmos, A. C. (2011). Ophiophagus hannah (King Cobra). Diet. Herpetological Review, 42, 297.

Tkach, V. V., Kuzmin, Y., & Brown, R. M. (2011). Rhabdias mcguirei sp. nov. (Nematoda: Rhabdiasidae) from the flying lizard, Draco spilopterus, (Squamata: Agamidae) of the northern Philippines. Acta Parasitologica, 56, 406-4111.

Brown, R. M., Siler, C. D., Oliveros, C. H., Diesmos, A. C., & Alcala, A. C. (2011). A new Gekko from Sibuyan Island, central Philippines. Herpetologica, 67, 460-476.

Brown, R. M., Diesmos, A. C., & Oliveros, C. (2011). A new flap-legged forest gecko (Genus Luperosaurus) from the northeastern Philippines. Journal of Herpetology, 45, 202-210.

Setiadi, M. I., McGuire, J. A., Brown, R. M., Zubairi, M., Iskandar, D. T., Andayani, N., Supriatna, J., & Evans, B. J. (2011). Adaptive radiation and ecological opportunity in Sulawesi and Philippine fanged frog (Limnonectes) communities. American Naturalist, 178, 221-240.

McLeod, D. S., Siler, C. D., Diesmos, A. C., Diesmos, M. L. D., Garcia, V. S., Arkonceo, A. O., Balaquit, K. L., Uy, C. C., Villaseran, M. M., Yarra, E. C., & Brown, R. M. (2011). Amphibians and Reptiles of Luzon Island, V: the herpetofauna of Angat Dam Watershed, Bulacan Province, Luzon Island, Philippines. Asian Herpetological Research, 2011, 177-198.

Siler, C. D., Welton, L. J., Siler, J. M., Brown, J., Bucol, A., Diesmos, A. C., & Brown, R. M. (2011). Amphibians and Reptiles, Luzon Island, Aurora Province and Aurora Memorial National Park, Northern Philippines: New island distribution records. Check List, 7, 182-195.

Siler, C. D., & Brown, R. M. (2011). Evidence for repeated acquisition and loss of complex body form characters in an insular clade of Southeast Asian semi-fossorial skinks. Evolution, 65, 2641–2663.

Swei, A., Rowley, J. J. L., Rödder, D., Diesmos, M. L. L., Diesmos, A. C., Briggs, C. J., Brown, R. M., Cao, T. T., Cheng, T. L., Chong, R. A., Han, B., Hero, J., Hoang, H. D., Kusrini, M. D., Le, D. T. T., McGuire, J. A., Meegaskumbura, M., Min, M., Mulcahy, D. G., Neang, T., Phimmachak, S., Rao, D., Reeder, N. M., Schoville, S. D., Sivongxay, N., Srei, N., Stöck, M., Stuart, B. L., Torres, L. S., Tran, D. T. A., Tunstall, T. S., Vietes, D., & Vredenburg, V. T. (2011). Is chytridiomycosis an emerging disease in Asia? PLoS ONE, 6, e23179. DOI:10.1371/journal.pone.0023179

Linkem, C. W., Diesmos, A. C., & Brown, R. M. (2011). Molecular systematics of the Philippine forest skinks (Reptilia: Scincidae: Sphenomorphus): testing morphological and biogeographic hypotheses of interspecific relationships. Zoological Journal of the Linnean Society, 163, 1217-1243.

Fuiten, A., Diesmos, A. C., Welton, L. J., Barley, A., Oberheide, B., Rico, E. L. B., & Brown, R. M. (2011). New species of stream frog from the mountains of Luzon Island, Philippines. Herpetologica, 67, 89-103.

Arifin, U., Iskandar, D. T., Bickford, D. P., Brown, R. M., Meier, R., & Kutti, S. N. (2011). Phylogenetic relationship within the genus Staurois (Amphibia, Ranidae) based on 16S rRNA sequences. Zootaxa, 2733, 39-52.

Hertwig, S. T., Das, I., Schweizer, M., Brown, R. M., & Haas, A. (2011). Phylogenetic relationships of the Rhacophorus everetti-group and implications for the evolution of reproductive modes in Philautus (Amphibia: Anura: Rhacophoridae). Zoologica Scripta, 41, 29-46.

Siler, C. D., Diesmos, A. C., Alcala, A. C., & Brown, R. M. (2011). Phylogeny of Philippine slender skinks (Scincidae: Brachymeles) reveals underestimated species diversity, complex biogeographical relationships, and cryptic patterns of lineage diversification. Molecular Phylogenetics and Evolution, 59, 53-65.

Siler, C. D., Fuitan, A. M., Jones, R. M., Alcala, A. C., & Brown, R. M. (2011). Phylogeny-based species delimitation in Philippine slender skinks (Reptilia: Squamata: Scincidae) II: taxonomic revision of Brachymeles samarensis and description of five new species. Herpetological Monographs, 25, 76-112.

Sanguila, M. B., Siler, C. D., Diesmos, A. C., Nuñeza, O., & Brown, R. M. (2011). Phylogeography and conservation implications of geographic structure of genetic variation and potential species boundaries in Philippine slender toads. Molecular Phylogenetics and Evolution, 61, 333-350.

Swei, A., Rowley, J. J. L., Rödder, D., Diesmos, M. L. L., Diesmos, A. C., Briggs, C. J., Brown, R. M., Cao, T. T., Cheng, T. L., Chong, R. A., Han, B., Hero, J., Hoang, H. D., Kusrini, M. D., Le, D. T. T., McGuire, J. A., Meegaskumbura, M., Min, M., Mulcahy, D. G., Neang, T., Phimmachak, S., Rao, D., Reeder, N. M., Schoville, S. D., Sivongxay, N., Srei, N., Stöck, M., Stuart, B. L., Torres, L. S., Tran, D. T. A., Tunstall, T. S., Vietes, D., & Vredenburg, V. T. (2011). Prevalence and distribution of chytridiomycosis throughout Asia. FrogLog, 98, 33-34.

Siler, C. D., Jones, R. M., Welton, L. J., & Brown, R. M. (2011). Redescription of tetradactyl, Philippine slender skinks (genus Brachymeles). Herpetologica, 67, 300-317.

Siler, C. D., Crombie, R. I., Diesmos, A. C., & Brown, R. M. (2011). Redescription of two poorly known loam-swimming skinks, Brachymeles bicolor and Brachymeles pathfinderi (Reptilia: Squamata: Scincidae) from the Philippines. Journal of Herpetology, 43, 355-369.

Esselstyn, J. A., Maher, S. P., & Brown, R. M. (2011). Species interactions during diversification and community assembly in an island radiation of shrews. PLoS ONE, 6, e21885 1–13.

Oliveros, C. H., Ota, H., Crombie, R. I., & Brown, R. M. (2011). The Herpetofauna of the Babuyan group of islands, northern Philippines (., & .). Scientific Publications of the Natural History Museum of the University of Kansas, 43, 1-20.

Holder, M. T., Sukumaran, J. A., & Brown, R. M. (2010). Bayesian Approaches to Phylogenetic Analysis. In D. K. Dey, S. Ghosh, & B. K. Mallick (Eds.), Bayesian Modeling in Bioinformatics (pp. 1-39). Chapman & Hall/CRC.

Brown, R. M., Diesmos, A. C., Duya, M. V., Garcia, H. J. D., & Rico, E. L. (2010). A new forest gecko (Squamata; Gekkonidae; Genus Luperosaurus) from Mt. Mantalingajan, southern Palawan Island, Philippines. Journal of Herpetology, 44, 37-48.

Siler, C. D., Balete, D. S., Diesmos, A. C., & Brown, R. M. (2010). A new legless loam-swimming lizard (Reptilia: Squamata: Scincidae: Genus Brachymeles) from the Bicol Peninsula, Luzon Island, Philippines. Copeia, 2010, 114-122.

Siler, C. D., Diesmos, A. C., & Brown, R. M. (2010). A new loam-swimming skink, genus Brachymeles (Reptilia: Squamata: Scincidae) from the Bicol faunal region, Luzon and Catanduanes islands, Philippines. Journal of Herpetology, 44, 49-60.

Linkem, C. A., Diesmos, A. C., & Brown, R. M. (2010). A new scincid lizard (Genus Sphenomorphus) from Palawan Island, Philippines. Herpetologica, 66, 67-79.

Linkem, C. W., Siler, C. D., Diesmos, A. C., & Brown, R. M. (2010). A new species of Gekko (Squamata: Gekkonidae) from central Luzon Island, Philippines. Zootaxa, 2396, 37-94.

Siler, C. D., Diesmos, A. C., Linkem, C. W., Diesmos, M., & Brown, R. M. (2010). A new species of limestone-forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae) from central Luzon Island, Philippines. Zootaxa, 2482, 49-63.

Welton, L. J., Siler, C. D., Bennet, D., Diesmos, A. C., Duya, M. R., Dugay, R., Rico, E. L., van Weerd, M., & Brown, R. M. (2010). A spectacular new Philippine monitor lizard reveals a hidden biogeographic boundary and a novel flagship species for conservation. Biology Letters, 6, 654–658.

Blackburn, D. C., Bickford, D. P., Diesmos, A. C., Iskandar, D. T., & Brown, R. M. (2010). An ancient origin for the enigmatic Flat-Headed Frogs (Bombinatoridae: Barbourula) from the islands of Southeast Asia. PLoS ONE, 5(8), e12090. DOI:10.1371/journal.pone.0012090

Rowley, J., Brown, R., Bain, R., Kusrini, M., Inger, R., Stuart, B., Wogan, G., Chan-ard, T., Trung, C. T., Diesmos, A. C., Iskandar, D. T., Lau, M., Ming, L. T., Makchai, S., Thy, N., Truong, N. Q., & Phimmachak, S. (2010). Impending conservation crisis for Southeast Asian amphibians. Biology Letters, 6, 336–338.

Esselstyn, J. A., Oliveros, C. H., Moyle, R. G., Peterson, A. T., McGuire, J. A., & Brown, R. M. (2010). Integrating phylogenetic and taxonomic evidence illuminates complex biogegraphic patterns along Huxley’s modification of Wallace’s Line. Journal of Biogeography, 37, 2054–2066. DOI:10.1111/j.1365-2699.2010.02378.x

Bauer, A. M., Parham, J. F., Brown, R. M., Stuart, B. L., Grismer, L. L., Papenfuss, T. J., Bohme, W., Savage, J., Carranza, S., Grismer, J., Wagner, P., Ananjeva, N., & Inger, R. F. (2010). On the availability of new Bayesian-delimited gecko names and the importance of character based definitions of species. Proceedings of the Royal Society of London B, 278, 490-492.

Welton, L. J., Siler, C., Diesmos, A. C., & Brown, R. M. (2010). Philippine bent-toed geckos of the Cyrtodactylus agusanensis complex: multilocus phylogeny, morphological diversity, and description of three new species. Herpetological Monographs, 24, 55-85.

Siler, C. D., Oaks, J. R., Esselstyn, J. A., Diesmos, A. C., & Brown, R. M. (2010). Phylogeny and biogeography of Philippine bent-toed geckos (Gekkonidae: Cyrtodactylus) contradict a prevailing model of Pleistocene diversification. Molecular Phylogenetics and Evolution, 55, 699-710.

Siler, C. D., & Brown, R. M. (2010). Phylogeny-based species delimitation in Philippine slender skinks (Reptilia: Squamata: Scincidae: Brachymeles): taxonomic revision of the pentadactyl species groups and description of three new species. Herpetological Monographs, 24, 1-54.

Welton, L. J., Siler, C., Diesmos, A. C., & Brown, R. M. (2010). Phylogeny-based species delimitation of southern Philippine bent-toed geckos and a new species of Cyrtodactylus (Squamata: Gekkonidae) from western Mindanao and the Sulu Archipelago. Zootaxa, 2390, 49-68.

Brown, R. M., Linkem, C. W., Siler, C. D., Sukumaran, J., Esselstyn, J. A., Diesmos, A. C., Iskandar, D. T., Bickford, D., Evans, B. J., McGuire, J. A., Grismer, L., Supriatna, J., & Andayani, N. (2010). Phylogeography and historical demography of Polypedates leucomystax in the islands of Indonesia and the Philippines: evidence for recent human-mediated range expansion? Molecular Phylogenetics and Evolution, 57, 598-619.

Linkem, C. A., Hesed, K., Diesmos, A. C., & Brown, R. M. (2010). Species boundaries and cryptic lineage diversity in a Philippine forest skink complex (Reptilia; Squamata; Scincidae: Lygosominae). Molecular Phylogenetics and Evolution, 56, 572–585.

Brown, R. M., Linkem, C. W., Balete, D., Duya, M. V., Diesmos, A. C., & Ferner, J. W. (2010). Species boundaries in Philippine montane forest skinks (Genus Sphenomorphus): three new species from the mountains of Luzon and clarification of the status of the poorly known S. beyeri, S. knollmanae, and S. laterimaculatus. Scientific Papers of the Natural History Museum of the University of Kansas, 42, 1-27.

Brown, R. M. (2009). Frogs in Island Archipelagos. In R. Gillespie & D. Clague (Eds.), Encyclopedia of Islands (pp. 347-351). Berkeley: University of California Press.

Brown, R. M., & Diesmos, A. C. (2009). Philippines, Biology. In R. Gillespie & D. Clague (Eds.), Encyclopedia of Islands (pp. 723-732). Berkeley: University of California Press.

Welton, L. J., Siler, C., Diesmos, A. C., & Brown, R. M. (2009). A new bent-toed gecko (Genus Cyrtodactylus) from southern Palawan Island, Philippines, and clarification of the taxonomic status of C. annulatus. Herpetologica, 65, 323–343.

Siler, C. D., Rico, E. L., Duya, M. R., & Brown, R. M. (2009). A new limb-reduced skink, genus Brachymeles (Reptilia: Squamata: Scincidae), from central Luzon Island, Philippines. Herpetologica, 65, 449–459.

Siler, C. D., McVay, J., Diesmos, A. C., & Brown, R. M. (2009). A new species of fanged frog (Dicroglossidae; genus Limnonectes) from southern Mindanao Island, Philippines. Herpetologica, 65, 105-114.

Siler, C. D., Diesmos, A. C., Alcala, A. C., & Brown, R. M. (2009). A new species of limestone forest frogs, genus Platymantis (Amphibia; Anura; Ceratobatrachidae) from Samar Island, Philippines. Herpetologica, 65, 92-104.

Esselstyn, J. A., Timm, R. M., & Brown, R. M. (2009). Do geological or climatic processes drive speciation in dynamic archipelagos? The evolution of shrew diversity on SE Asian islands. Evolution, 63, 2595–2610. DOI:10.1111/j.1558-5646.2009.00743.x

Wiens, J. J., Sukumaran, J. S., Pyron, R. A., & Brown, R. M. (2009). Evolutionary and biogeographic origins of high tropical diversity in old world frogs (Ranidae). Evolution, 64, 1217–1231. DOI:10.1111/j.1558-5646.2009.00610.x

Setiadi, M. I., Hamidy, A., Abidin, Z., Brown, R. M., Peterson, A. T., Li, X., & Evans, B. J. (2009). Geographic genetic structure of herpetofauna on Halmahera Island, Indonesia: implications for the Aketajawe-Lolobata National Park. Conservation Biology, 24(553-562).

Matsui, M., Tominaga, A., Liu, W., Khonsue, W., Grismer, L. L., Diesmos, A. C., Das, I., Sudin, A., Yambun, P., Yong, H., Sukumaran, J., & Brown, R. M. (2009). Phylogenetic Relationships of Ansonia from Southeast Asia inferred from mitochondrial DNA sequences: Systematic and Biogeographic Implications (Anura: Bufonidae). Molecular Phylogenetics and Evolution, 54, 561-570. DOI:10.1016/j.ympev.2009.08.003

Brown, R. M., Oliveros, C., Siler, C. D., & Diesmos, A. C. (2009). Phylogeny of Gekko from the northern Philippines, and description of a new species from Calayan Island. Journal of Herpetology, 43, 620-635.

Brown, R. M., Siler, C. D., Diesmos, A. C., & Alcala, A. C. (2009). The Philippine frogs of the genus Leptobrachium (Anura; Megophryidae): phylogeny-based species delimitation, taxonomic revision, and descriptions of three new species. Herpetological Monographs, 23, 1-44.

Esselstyn, J. A., & Brown, R. M. (2009). The role of repeated sea-level fluctuations in the generation of shrew (Soricidae: Crocidura) diversity in the Philippine Archipelago. Molecular Phylogenetics and Evolution, 53, 171–181.

Bain, R., Biju, S. D., Brown, R., Das, I., Diesmos, A., Dutta, S., Gower, D., Inger, R., Iskandar, D., Kaneko, Y., Lau, M. W. N., Meegaskumbura, M., Ohler, A., Papenfuss, T., Pethiyagoda, R., Stuart, B., Wilkinson, M., & Xie, F. (2008). Amphibians of the Indomalayan Realm. In S. N. Stuart, M. Hoffmann, J. S. Chanson, N. A. Cox, R. Berridge, P. Ramani, & B. E. Young (Eds.), Threatened Amphibians of the World (pp. 74-79). Barcelona, Spain; Gland, Switzerland; and Arlington, VA, USA: Lynx Ediciones, IUCN - The World Conservation Union, and Conservation International.

Ron, S., & Brown, R. M. (2008). Filling the black hole: challenges in taxonomy to protect amphibians. In S. N. Stuart, M. Hoffmann, J. S. Chanson, N. A. Cox, R. Berridge, P. Ramani, & B. E. Young (Eds.), Threatened Amphibians of the World (pp. 133). Barcelona, Spain; Gland, Switzerland; and Arlington, VA, USA: Lynx Ediciones, IUCN - The World Conservation Union, and Conservation International.

Bickford, D. P., Supriatna, J., Andayani, Y., Iskandar, D., Evans, B. J., Brown, R. M., Townsend, T., Azhari, D., & McGuire, J. (2008). Indonesia’s protected areas need more protection: suggestions from island examples. In N. S. Sodhi, G. Acciaioli, M. Erb, & A. K.-J. Tan (Eds.), Biodiversity and Human Livelihoods in Protected Areas: Case Studies from the Malay Archipelago (pp. 53-77). Cambridge, UK: Cambridge University Press.

Brown, R. M., Diesmos, A. C., & Alcala, A. C. (2008). Philippine amphibian biodiversity is increasing in leaps and bounds. In S. N. Stuart, M. Hoffmann, J. S. Chanson, N. A. Cox, R. Berridge, P. Ramani, & B. E. Young (Eds.), Threatened Amphibians of the World (pp. 82-83). Barcelona, Spain; Gland, Switzerland; and Arlington, VA, USA: Lynx Ediciones, IUCN - The World Conservation Union, and Conservation International.

Brown, R. M. (2008). [Review of the book Field Guide to the Amphibians and Reptiles of Bali, J. L. McKay (Kreiger Publishing, FL)]. Quarterly Review of Biology, 83, 122–123.

Brown, R. M. (2008). The Frogs of the Solomon Islands (P. Pikacha, C. Morrison, & S. Richards). Institute of Applied Science, University of the South Pacific.

Evans, B. J., McGuire, J. A., Brown, R. M., Andayani, N., & Supriatna, J. (2008). A coalescent framework for comparing alternative models of population structure with genetic data: evolution of Celebes toads. Biology Letters, 4, 430-433.

Brown, R. M., Oliveros, C., Siler, C. D., & Diesmos, A. C. (2008). A new Gekko from the Babuyan Islands, northern Philippines. Herpetologica, 64, 305–320.

Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, M. I., Bickford, D., & Brown, R. M. (2008). A new species of bent-toe gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, eastern Indonesia. Herpetologica, 64, 232-242.

Hayden, C. J., Brown, R. M., Gillespie, G., Setiadi, M. I., Linkem, C. W., Iskandar, D. T., Bickford, D. P., Riyanto, A., Mumpuni, I., & McGuire, J. A. (2008). A new species of bent-toed gecko Cyrtodactylus Gray, 1827, (Squamata: Gekkonidae) from the island of Sulawesi, Indonesia. Herpetologica, 64, 109-120.

Lim, B. K., Edger, J. L., Peterson, A. T., Robbins, M. B., Clayton, D. H., Bush, S. E., & Brown, R. M. (2008). Biodiversity in China: Lost in the Masses? Harvard Asia Quarterly, 11, 12-23.

Diesmos, A. C., Brown, R. M., Alcala, A. C., & Sison, R. V. (2008). Status and Distribution of Nonmarine Turtles of the Philippines. Chelonian Conservation and Biology, 7, 157–177.

Brown, R. M., & Richards, S. J. (2008). Two new frogs of the genus Platymantis (Anura: Ceratobatrachidae) from the Isabel Island group, Solomon Islands. Zootaxa, 1888, 47-68.

Parra, G., Brown, R., Hanken, J., Hedges, B., Heyer, R., Kuzmin, S., Lavilla, E., Lötters, S., Pimenta, B., Richards, S., Rödel, M. O., de Sá, R. O., & Wake, D. (2007). Systematics and Conservation. In C. Gascon, J. P. Collins, R. D. Moore, D. R. Church, J. E. McKay, & J. R. Mendelson (Eds.), Amphibian Conservation Action Plan (pp. 45-48). Gland, Switzerland and Cambridge, UK: IUCN/SSC Amphibian Specialist Group.

Brown, R. M. (2007). Introduction to Robert F. Inger’s Systematics and Zoogeography of Philippine Amphibia. Systematics and Zoogeography of Philippine Amphibia, 1-17. Kota Kinabalu: Natural History Publications.

Iverson, J. B., Brown, R. M., Akre, T. S., Near, T. J., Le, M., Thomson, R. C., & Starkey, D. E. (2007). In search of the tree of life for turtles. In H. B. Shaffer, N. N. FitzSimmons, A. Georges, & A. Rhodin (Eds.), Chelonian Research Monographs 4. Defining Turtle Diversity: Proceedings of a Workshop on Genetics, Ethics and Taxonomy of Fresh Water Turtles and Tortoises (pp. 85–106).

Lehn, C., Das, I., Forstner, M. R. J., & Brown, R. M. (2007). Responsible vouchering in turtle research: an introduction and recommendations. In H. B. Shaffer, N. N. FitzSimmons, A. Georges, & A. Rhodin (Eds.), Chelonian Research Monographs 4. Defining Turtle Diversity: Proceedings of a Workshop on Genetics, Ethics and Taxonomy of Fresh Water Turtles and Tortoises (pp. 147–156).

Brown, R. M., & Gonzalez, J. C. (2007). A new forest frog of the genus Platymantis (Amphibia; Anura: Ranidae) from the Bicol Peninsula of Luzon Island, Philippines. Copeia, 2007, 251-266.

Brown, R. M., Diesmos, A. C., & Duya, M. V. (2007). A new species of Luperosaurus (Squamata: Gekkonidae) from the Sierra Madre mountain range of northern Luzon Island, Philippines. Raffles Bulletin of Zoology, 55, 153–160.

Gaulke, M., Rösler, H., & Brown, R. M. (2007). A new species of Luperosaurus (Squamata; Gekkonidae) from Panay Island, Philippines, with comments on the taxonomic status of Luperosaurus cumingii (Gray, 1845). Copeia, 2007, 413-425.

Wallach, V., Brown, R. M., Diesmos, A. C., & Gee, G. V. A. (2007). An enigmatic new species of blind snake from Luzon Island, northern Philippines (Serpentes: Typhlopidae). Journal of Herpetology, 41, 690–702.

Dudley, R., Byrnes, G., Yanoviak, S., Borrell, B., Brown, R. M., & McGuire, J. (2007). Gliding and the functional origins of flight: biomechanical novelty or necessity? Annual Review of Ecology, Evolution, and Systematics, 38, 179–201.

Castoe, T. A., Smith, E. N., Brown, R. M., & Parkinson, C. L. (2007). Higher-level phylogeny of Asian and American coralsnakes, their placement within the Elapidae (Squamata), and the systematic affinities of the enigmatic Asian coralsnake Hemibungarus calligaster (Wiegmann, 1834). Zoological Journal of the Linnean Society, 151, 809–831.

McGuire, J. A., Brown, R. M., Riyanto, A., & Andayani, N. (2007). The flying lizards of the Draco lineatus group (Squamata: Iguania: Agamidae): A taxonomic revision with descriptions of two new species. Herpetological Monographs, 21, 180-213.

Brown, R. M. (2006). Recognition of Philippine amphibian diversity. In C. S. Catibog-Sinha & L. R. Heaney (Eds.), Philippine Biodiversity: Principles and Practice (pp. 45). Quezon City: Haribon Foundation for Conservation of Natural Resources.

Brown, R. M. (2006). Species concepts. In C. S. Catibog-Sinha & L. R. Heaney (Eds.), Philippine Biodiversity: Principles and Practice (pp. 43-44). Quezon City: Haribon Foundation for Conservation of Natural Resources.

Brown, R. M. (2006). The use of genetic data for inferring species limits. In C. S. Catibog-Sinha & L. R. Heaney (Eds.), Philippine Biodiversity: Principles and Practice (pp. 87–90). Quezon City: Haribon Foundation for Conservation of Natural Resources.

Rösler, H., Siler, C. D., Brown, R. M., Demegillo, A. D., & Gaulke, M. (2006). Gekko ernstkelleri sp. n.—a new gekkonid lizard from Panay Island, Philippines. Salamandra, 42, 197–211.

Brown, R. M. (2006). A case of suspected coral snake mimicry by lepidopteran larvae. Raffles Bulletin of Zoology, 54, 225–227.

Brown, R. M., Richards, S. J., Sukumaran, J., & Foufopoulos, J. (2006). A new morphologically cryptic species of forest frog (genus Platymantis) from New Britain Island, Bismarck Archipelago. Zootaxa, 1334, 45-68.

Bossuyt, F., Brown, R. M., Hillis, D. M., Cannatella, D. C., & Milinkovitch, M. C. (2006). Late Cretaceous diversification resulted in continent-scale regionalism in the cosmopolitan frog family Ranidae. Systematic Biology, 55, 579–594.

Brown, R. M., Foufopoulos, J., & Richards, S. J. (2006). New species of Platymantis (Amphibia; Anura: Ranidae) from New Britain and redescription of the poorly known Platymantis nexipus. Copeia, 2006, 674–695.

Diesmos, A. C., Diesmos, M. L., & Brown, R. M. (2006). Status and distribution of alien invasive frogs in the Philippines. Journal of Environmental Science and Management, Philippines, 9, 41–53.

Diesmos, A. C., Parham, J. F., Stuart, B. L., & Brown, R. M. (2005). The phylogenetic position of the recently rediscovered Philippine forest turtle (Bataguridae: Heosemys leytensis). Proceedings of the California Academy of Sciences, 56, 31-41.

Foufopoulos, J., & Brown, R. M. (2004). A new frog of the genus Platymantis (Amphibia; Anura; Ranidae) from New Britain,with a redescription of the poorly-known Platymantis macrosceles. Copeia, 2004, 825–841.

Diesmos, A. C., Brown, R. M., & Gee, G. V. A. (2004). Preliminary report on the amphibians and reptiles of Balbalasang-Balbalan National Park, Luzon Island, Philippines. Sylvatrop, Technical Journal of Philippine Ecosystems and Natural Resources, 13, 63-80.

Diesmos, A. C., Gee, G. V. A., Brown, R. M., Widmann, P. J., & Dimalibot, J. C. (2004). Rediscovery of the Philippine forest turtle, Heosemys leytensis (Chelonia; Bataguridae), from Palawan Island, Philippines. Asiatic Herpetological Research, 10, 22-27.

Evans, B. J., Brown, R. M., McGuire, J. A., Supriatna, J., Andayani, N., Diesmos, A. C., Iskandar, D., Melnick, D. J., & Cannatella, D. C. (2003). Phylogenetics of fanged frogs: testing biogeographical hypotheses at the interface of the Asian and Australian faunal zones. Systematic Biology, 52, 794–819.

Brown, R. M., Dolino, C. N., Alcala, E., Diesmos, A. C., & Alcala, A. C. (2003). The advertisement calls of two endangered species of endemic Philippine frogs: Platymantis spelaeus and P. insulatus (Anura; Ranidae). The Silliman Journal, 43, 91–109.

Brown, R. M., Diesmos, A. D., & Fernandez, R. B. (2003). The rising costs of cabbage and potatoes. Haring Ibon, 9, 8-12.

Diesmos, A. C., Brown, R. M., Alcala, A. C., Sison, R. V., Afuang, L. E., & Gee, G. V. A. (2002). Philippine amphibians and reptiles. In P. S. Ong, L. E. Afuang, & R. G. Rosell-Ambal (Eds.), Philippine Biodiversity Conservation Priorities: a Second Iteration of the National Biodiversity Strategy and Action Plan (pp. 26–44). Quezon City, Philippines: Department of the Environment and Natural Resources–Protected Areas and Wildlife Bureau, Conservation International Philippines, Biodiversity Conservation Program–University of the Philippines Center for Integrative and Developmental Studies, and Foundation for the Philippine Environment.

Diesmos, A. C., Brown, R. M., & Alcala, A. C. (2002). A new species of narrow-mouthed frog (Amphibia; Anura; Microhylidae: genus Kaloula) from the mountains of southern Luzon and Polillo islands, Philippines. Copeia, 2002, 1037–1051.

Brown, R. M., & Diesmos, A. C. (2002). Application of lineage-based species concepts to oceanic island frog populations: the effects of differing taxonomic philosophies on the estimation of Philippine biodiversity. The Silliman Journal, 42, 133–162.

Brown, R. M., & Guttman, S. I. (2002). Phylogenetic systematics of the Rana signata complex of Philippine and Bornean stream frogs: reconsideration of Huxley’s modification of Wallace’s Line at the Oriental-Australian faunal zone interface. Biological Journal of the Linnaean Society, 76, 393–461.

Brown, R. M., Diesmos, A. C., & Alcala, A. C. (2002). The state of Philippine herpetology and the challenges for the next decade. The Silliman Journal, 42, 18-87.

Brown, R. M., Fernandez, R., Rivero, C., Buenviaje, R., & Diesmos, A. (2002). Mt. Isarog's herpetological wonders. Haring Ibon, 3, 12-16.

Infante, C. R., Brown, R. M., Diesmos, A. C., Gee, G. V. A., & Dimalibot, J. (2002). The search for the lost treasure of Palawan: the Philippine flat-headed frog. Haring Ibon, 8, 12–15.

Brown, R. M., Leviton, A. E., Ferner, J. W., & Sison, R. V. (2001). A new species of snake in the genus Hologerrhum (Reptilia; Squamata; Serpentes) from Panay Island, Philippines. Asiatic Herpetological Research, 9, 9-22.

Ferner, J. W., Brown, R. M., Sison, R. V., & Kennedy, R. S. (2001). The amphibians and reptiles of Panay Island, Philippines. Asiatic Herpetological Research, 9, 34-70.

Brown, R. M., McGuire, J. A., Ferner, J. W., Icarangal, N., & Kennedy, R. S. (2000). Amphibians and reptiles of Luzon Island, II: preliminary report on the herpetofauna of Aurora Memorial National Park, Philippines. Hamadryad, 25, 175–195.

Brown, R. M., Supriatna, J., & Ota, H. (2000). Discovery of a new species of Luperosaurus (Squamata; Gekkonidae) from Sulawesi, with a phylogenetic analysis of the genus and comments on the status of L. serraticaudus. Copeia, 2000, 191-209.

Brown, R. M., & Iskandar, D. T. (2000). Nest site selection, larval hatching, and advertisement calls of Rana arathooni (Amphibia; Anura; Ranidae) from southwestern Sulawesi (Celebes) Island, Indonesia. Journal of Herpetology, 34, 404-413.

Brown, R. M., McGuire, J. A., & Diesmos, A. C. (2000). Status of some Philippine frogs related to Rana everetti (Anura: Ranidae), description of a new species, and resurrection of Rana igorota Taylor 1922. Herpetologica, 56, 81-104.

Brown, R. M., & Diesmos, A. C. (2000). The lizard genus Luperosaurus: taxonomy, history, and conservation prospects for some of the world's rarest lizards. Sylvatrop: Technical Journal of Philippine Ecosystems and Natural Resources, 10, 107–124.

Heaney, L. R., Diesmos, A., Tabaranza, B., Mallari, A., Brown, R., & Gee, G. (2000). Beacon of hope: a first report from Kalinga Province in the northern Central Cordillera. Haring Ibon, 2, 14-18.

Brown, R. M., & Alcala, A. C. (2000). Geckos, cave frogs, and small land-bridge islands in the Visayan sea. Haring Ibon, 2, 19-22.

Brown, R. M., McGuire, J. A., Ferner, J. W., & Alcala, A. C. (1999). A new diminutive species of skink (Squamata; Scincidae; Lygosominae: Sphenomorphus) from Luzon Island, Republic of the Philippines. Copeia, 1999, 362–370.

Brown, R. M., Leviton, A. E., & Sison, R. V. (1999). Description of a new species of Pseudorabdion from Panay Island, Philippines, with a revised key to the genus. Asiatic Herpetological Research, 8, 7-12.

Brown, R. M. (1999). New species of parachute gecko (Squamata; Gekkonidae; Genus Ptychozoon) from northeastern Thailand and Central Vietnam. Copeia, 1999, 990–1001.

Brown, W. C., Alcala, A. C., & Brown, R. M. (1998). Taxonomic status of Cornufer worcesteri. Journal of Herpetology, 33, 131-133.

Ferner, J. W., Brown, R. M., & Greer, A. E. (1997). A new genus and species of closed canopy forest skinks from the Philippine islands. Journal of Herpetology, 31, 187–192.

Brown, R. M., Ferner, J. W., & Diesmos, A. C. (1997). Definition of the Philippine parachute gecko, Ptychozoon intermedium Taylor 1915 (Reptilia: Squamata: Gekkonidae): redescription, designation of a neotype, and comparisons with related species. Herpetologica, 53, 357–373.

Brown, W. C., Brown, R. M., Alcala, A. C., & Frost, D. (1997). Replacement name for Platymantis reticulatus Brown, Brown, and Alcala, 1997 (Ranidae: Raninae). Herpetological Review, 28, 131.

Brown, W. C., Brown, R. M., & Alcala, A. C. (1997). Species of the hazelae group of Platymantis from the Philippines with descriptions of two new species. Proceedings of the California Academy of Sciences, 49, 405–421.

Brown, R. M., Ferner, J. W., Sison, R. V., Gonzales, P. C., & Kennedy, R. S. (1996). Amphibians and reptiles of the Zambales mountains of Luzon Island, Philippines. Herpetological Natural History, 4, 1-17.

Brown, R. M., Ferner, J. F., & Ruedas, L. A. (1995). A new species of lygosomine lizard (Reptilia; Lacertilia; Scincidae; Sphenomorphus) from Mt. Isarog, Luzon Island, Philippines. Proceedings of the Biological Society of Washington, 108, 18-28.

Brown, R. M., & Taylor, D. H. (1995). Compensatory escape mode trade-offs between locomotor performance and maneuvering behavior across larval ontogeny of the wood frog Rana sylvatica. Copeia, 1995, 1-7.

Brown, R. M., Taylor, D. H., & Gist, D. H. (1995). Effects of caudal autotomy on locomotor performance of wall lizards (Podarcis muralis). Journal of Herpetology, 29, 98–105.

Brown, R. M., Ferner, J. F., & Sison, R. V. (1995). Rediscovery and redescription of Sphenomorphus beyeri (Lacertilia; Scincidae) from the Zambales Mountains of Luzon, Philippines. Proceedings of the Biological Society of Washington, 108, 6-17.

Brown, R. M., Gist, D. H., & Taylor, D. H. (1995). The home range ecology of an introduced population of wall lizard (Podarcis muralis) in Cincinnati, Ohio. American Midland Naturalist, 133, 186–202.

Selected Grants

Brown, R., Describing Cryptic Species Diversity in Philippine Forest Frogs, Conservation International Threatened Species Fund Grant, $5,000, Not-for-Profit (not Foundation). Status: Funded.

Brown, R., (Principal), Robbins, M., (Co-Principal), Webster, M., (Principal), Scholes, E., (Co-Principal), Bostwick, K., (Co-Principal), Friel, J., (Co-Principal), Budney, G., (Co-Principal), Centralized Digital Archive of Vouchered Animal Communication Signals Source of Support, National Science Foundation, Digitization ADBC Program Grant, $198,000, 09/01/2012 - 08/31/2016. Federal. Status: Funded.

Brown, R., (Principal), Grismer, J., (Co-Principal), Revisiting Przhevalsky's Expeditions: Population Genetics in The Gobi Desert, NGS 9535-14, National Geographic Society, $22,468, 01/09/2015 - 01/09/2016. Not-for-Profit (not Foundation). Status: Funded.

Brown, R., (Principal), Moye, R., (Co-Principal), Peterson, T., (Co-Principal), Re-organization of Philippine Rainforest Biodiversity Following Disturbance on Regional Scales from a Powerful Typhoon, National Science Foundation, Systematic Panel Proposal: RAPID, $125,000, 01/05/2014 - 01/05/2015. Federal. Status: Funded.

Brown, R., (Principal), Moyle, R., (Co-Principal), Clayon, D., (Co-Principal), Bush, S., (Co-Principal), A Comprehensive Biotic Survey of Philippine Land Vertebrates and Their Parasites, DEB 0743491, National Science Foundation, Biotic Surveys and Inventories Panel, $900,942, 01/01/2008 - 12/31/2013. Federal. Status: Funded.

Vredenburg, V., (Principal), Brown, R., (Co-Principal), Can we prevent a chytridiomycosis epidemic in the Philippines?, USFS 96200-1-G272, United States Forest Service, Wildlife Without Borders - Amphibians in Decline program, $48,961, 01/01/2011 - 01/31/2013. Federal. Status: Funded.

Brown, , (Principal), Holder, , (Co-Principal), Oaks, , (Co-Principal), Comparative Phylogeography of a Dynamic Archipelago, DEB 1011423, National Science Foundation, Dissertation Improvement Grant Panel, $14,886, 01/01/2010 - 12/31/2012. Federal. Status: Funded.

Brown, , (Principal), Linkem, C., (Co-Principal), Integrative Morphological Evolution in Philippine Forest Lizards of the Genus Sphenomorphus, DEB-0910341, National Science Foundation, Dissertation Improvement Grant Panel, $14,800, 01/01/2009 - 12/31/2011. Federal. Status: Funded.

Moyle, R., Brown, R., Revisiting community assembly on Islands integrating ecological and evolutionary drivers within bird, frog, and skink communities across nested tropical Pacific scales, General Research Fund (University of Kansas), $14,310, 03/30/2011. University (KU or KUMC). Status: Funded.

Brown, R., (Principal), McGuire, J., (Principal), Evans, B., (Principal), Collaborative Research: Comparative Biogeography of Sulawesi—Phylogenetic and Coalescent Analyses of Diversification in Frogs, Lizards, and Monkeys, DEB 0640737, National Science Foundation, Systematic Panel Grant, $244,481, 01/01/2007 - 12/31/2010. Federal. Status: Funded.

Brown, R., (Principal), Weghorst, J., (Co-Principal), Shekelle, M., (Co-Principal), Neri-Arboleda, I., (Co-Principal), Diesmos, M., (Co-Principal), Duya, L., (Co-Principal), Establishing Conservation Priorities for Philippine Tarsiers (Tarsius syrichta) Using Survey, Population Census, Bioacoustic, and Molecular Conservation Genetic Techniques to Identify Evolutionary Significant Units for Conservation, NGS 8446-08, National Geograhic Society, $24,605, 08/01/2007 - 08/31/2010. Not-for-Profit (not Foundation). Status: Funded.

Moyle, R., Brown, R., Facilities upgrades for the Dyche Hall Molecular Phylogenetics Lab, College of Liberal Arts and Sciences Instructional Technology Fund Grant, $8,200, 02/28/2010. University (KU or KUMC). Status: Funded.

Brown, R., Holder, M., Moyle, R., Trueb, L., Peterson, T., Wiley, E., The Dyche Hall Computational Lab Teaching Facility, Phase III, College of Liberal Arts and Sciences Instructional Technology Fund Grant, $7,000, 12/31/2009. University (KU or KUMC). Status: Funded.

Brown, , (Principal), Siler, C., (Co-Principal), Historical Processes and Genetic Implications of Limb Reduction and Loss in an Island Skink Lineage, DEB 0804115, National Science Foundation, Dissertation Improvement Grant Panel, $11,950, 01/01/2008 - 12/31/2009. Federal. Status: Funded.

Peterson, T., (Principal), Brown, R., (Co-Principal), Clayton, D., (Co-Principal), Lim, B., (Co-Principal), Biodiversity Surveys in the Southern Borderlands of the People’s Republic of China, DEB 0344430, National Science Foundation, Biological Surveys and Inventories Grant, $750,000, 01/01/2004 - 12/31/2009. Federal. Status: Funded.

Brown, R., Sasa, Mahmood, Amphibian Productivity in Costa Rica's Dry Forest: Implications for Wetlands Protection as Reproductive Sites, University of Kansas International Programs Costa Rica / KU Collaboration Fund Grant, $10,000, 12/31/2008. University (KU or KUMC). Status: Funded.

Trueb, L., (Principal), Brown, R., (Co-Principal), AmphibiaTree—an Integrated, Phylogenetic and Bioinformatic Approach to the Tree of Amphibians, EF-0334952, National Science Foundation Assembling the Tree Of Life Grant, $237,000, 01/01/2003 - 12/31/2008. Federal. Status: Funded.

Brown, R., Moyle, R., Robbins, M., Peterson, T., Solid-State Digital Recording Equipment for Teaching and Research Applications in Ornithology and Herpetology, College of Liberal Arts and Sciences Instructional Technology Fund Grant, $19,440, 01/31/2008. University (KU or KUMC). Status: Funded.

Brown, R., Moyle, R., Trueb, L., Peterson, T., Wiley, E., The Dyche Hall Computational Lab Teaching Facility, Phase II, College of Liberal Arts and Sciences Instructional Technology Fund Grant, $19,083, 01/31/2007. University (KU or KUMC). Status: Funded.

Brown, R., Stages of Evolutionary Diversification in Replicated Radiations of Forest Frogs from SE Asia and the SW Pacific, General Research Fund (University of Kansas), $11,659, 02/28/2006. University (KU or KUMC). Status: Funded.

Brown, R., Peterson, T., Wiley, E., Trueb, L., A New Computational Laboratory and Teaching Facility for Dyche Hall, College of Liberal Arts and Sciences Instructional Technology Fund Grant, $19,177, 01/31/2006. University (KU or KUMC). Status: Funded.


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